As the Wanteds, Unwanteds, and Necessaries struggle to adjust to changes in their society, Mr. Today begins training fourteen-year-old Alex to replace him as Artime's leader one day while Alex's disgraced twin, Aaron, connives to take over Quill.
The summer after her first year of college, Isobel "Belly" Conklin is faced with a choice between Jeremiah and Conrad Fisher, brothers she has always loved, when Jeremiah proposes marriage and Conrad confesses that he still loves her.